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1、Chapter1Passage1HumanBodyInthispassageyouwilllearn:1.Classificationoforgansystems2.Structureandfunctionofeachorgansystem3.AssociatedmedicaltermsTounderstandthehumanbodyitisnecessarytounderstandhowitspartsareputtogetherandhowtheyfunction.Thestudyofthebodysstructureiscalledanatomy;thestudyofthebodysfu
2、nctionisknownasphysiology.Otherstudiesofhumanbodyincludebiology,cytology,embryology,histology,endocrinology,hematology,immunology,psychologyetc.了解人體各部分的組成及其功能,對于認(rèn)識人體是必需的。研究人體結(jié)構(gòu)的科學(xué)叫解剖學(xué);研究人體功能的科學(xué)叫生理學(xué)。其他研究人體的科學(xué)包括生物學(xué)、細(xì)胞學(xué)、胚胎學(xué)、組織學(xué)、內(nèi)分泌學(xué)、血液學(xué)、遺傳學(xué)、免疫學(xué)、心理學(xué)等等。Anatomistsfinditusefultodividethehumanbodyintotensys
3、tems,thatis,theskeletalsystem,themuscularsystem,thecirculatorysystem,therespiratorysystem,thedigestivesystem,theurinarysystem,theendocrinesystem,thenervoussystem,thereproductivesystemandtheskin.Theprincipalpartsofeachofthesesystemsaredescribedinthisarticle.解剖學(xué)家發(fā)現(xiàn)把整個人體分成骨骼、肌肉、循環(huán)、呼吸、消化、泌尿、內(nèi)分泌、神經(jīng)、生殖系統(tǒng)以
4、及感覺器官的做法是很有幫助的。本文描繪并闡述了各系統(tǒng)的主要部分。Theskeletalsystemismadeofbones,jointsbetweenbones,andcartilage.Itsfunctionistoprovidesupportandprotectionforthesofttissuesandtheorgansofthebodyandtoprovidepointsofattachmentforthemusclesthatmovethebody.Thereare206bonesinthehumanskeleton.Theyhavevariousshapes-long,shor
5、t,cube-shaped,flat,andirregular.Manyofthelongboneshaveaninteriorspacethatisfilledwithbonemarrow,wherebloodcellsaremade.骨骼系統(tǒng)由骨、關(guān)節(jié)以及軟骨組成。它對軟組織及人體器官起到支持和保護(hù)作用,并牽動骨胳肌,引起各種運(yùn)動。人體有206根骨頭。骨形態(tài)不一,有長的、短、立方的、扁的及不規(guī)則的。許多長骨里有一個內(nèi)層間隙,里面充填著骨髓,這即是血細(xì)胞的制造場所。Ajointiswherebonesarejoinedtogether.Theconnectioncanbesoclosetha
6、tnomovementispossible,asisthecaseintheskull.Otherkindsofjointspermitmovement:eitherbackandforthinoneplane-aswiththehingejointoftheelbow-ormovementaroundasingleaxis-aswiththepivotjointthatpermitstheheadtorotate.Awiderangeofmovementispossiblewhentheball-shapedendofonebonefitsintoasocketattheendofanoth
7、erbone,astheydointheshoulderandhipjoints.關(guān)節(jié)把骨與骨連接起來。顱骨不能運(yùn)動,是由于骨與骨之間的連接太緊密。但其它的關(guān)節(jié)可允許活動,如一個平面上的前后屈伸運(yùn)動,如肘關(guān)節(jié);或是繞軸心旋轉(zhuǎn)運(yùn)動,如樞軸點(diǎn)允許頭部轉(zhuǎn)動。如果一根骨的球形末端插入另一根骨的臼槽里,大輻度的運(yùn)動(如肩關(guān)節(jié)、髖關(guān)節(jié))即成為可能。Cartilageisamoreflexiblematerialthanbone.Itservesasaprotective,cushioninglayerwherebonescometogether.Italsoconnectstheribstothebrea
8、stboneandprovidesastructuralbaseforthenoseandtheexternalear.Aninfantsskeletonismadeofcartilagethatisgraduallyreplacedbyboneastheinfantgrowsintoanadult.軟骨是一種比一般骨更具韌性的物質(zhì)。它是骨連結(jié)的保護(hù)、緩沖層。它把肋骨與胸骨連結(jié)起來,也是鼻腔與內(nèi)耳的結(jié)構(gòu)基礎(chǔ)。一個嬰兒的骨骼就是由軟骨組成,然后不斷生長、骨化,使嬰兒長大成人。Themuscularsystemallowsthebodytomove,anditscontractionsproduc
9、eheat,whichhelpsmaintainaconstantbodytemperature.Striatedmusclescanbeconsciouslycontrolled.Theendsofthesemusclesareattachedtodifferentbonesbyconnectivetissuebandssothatwhenthemusclecontracts,onebonemovesinrelationtotheother.Thismakesitpossibletomovethewholebody,aswhenwalking,ortomovejustonepartofthe
10、body,aswhenbendingafinger.Contractionsoftheheartandsmoothmusclesarenotunderconsciouscontrol.Smoothmusclesarefoundinthewallsoforganssuchasthestomachandtheintestinesandservetomovethecontentsoftheseorgansthroughthebody肌肉系統(tǒng)使軀體運(yùn)動,肌肉收縮產(chǎn)生的熱有助于維持一個恒定的體溫。人體能夠有意識地控制條紋肌。結(jié)締組織使肌肉末端附著于不同的骨面上,所以當(dāng)肌肉收縮時,兩骨彼此靠近而產(chǎn)生運(yùn)動。
11、這也就使整個人體可以運(yùn)動起來,如走路,運(yùn)動軀體某個部位,如彎曲手指。心臟收縮和平滑肌收縮就不是被意識所控制的。器官壁內(nèi)層的平滑肌,如胃腸壁的平滑肌把胃腸中的物質(zhì)運(yùn)送到全身。Thecirculatorysystem.Allpartsofthebodymusthavenourishmentandoxygeninordertofunctionandgrow,andtheirwasteproductsmustberemovedbeforetheyaccumulateandpoisonthebody.Thecirculatorysystemdistributesneededmaterialsandrem
12、ovesunneededones.Itismadeupoftheheart,bloodvessels,andblood,whichtogethermakeupthecardiovascularsystem.Thebloodisalsopartofthebodysdefensesystem.Ithasantibodiesandwhitebloodcellsthatprotectthebodyagainstforeigninvaders.循環(huán)系統(tǒng):機(jī)體的所有部分需要營養(yǎng)物質(zhì)和氧氣來使之發(fā)揮功能和生長,也需要在這些器官所產(chǎn)生的廢物積聚而危害生命之前將其排除。循環(huán)系統(tǒng)運(yùn)送有用物質(zhì),排泄廢物。心血管系統(tǒng)
13、是循環(huán)系統(tǒng)的組成之一;心血管系統(tǒng)包括心臟、血管及血液。血液也是機(jī)體防御系統(tǒng)的一個部分,血液中有抗體及白細(xì)胞來防止機(jī)體受到外來的侵襲。Theheartisamusclethatisdividedintotwonearlyidenticalhalves:onehalfreceivesbloodfromthelungsandsendsittotherestofthebody,theotherhalfsendsbloodthathastraveledthroughthebodybacktothelungs.Whentheheartmusclecontracts,thebloodisforcedouti
14、ntoarteriesandenterssmallcapillaries.Bloodreturnstotheheartthroughveins.心臟是一塊被分為幾乎對等兩半的肌肉。一半吸收來自肺部的血液,并將血液運(yùn)送到機(jī)體的其余部位,另一半使流經(jīng)全身的血液回流人肺。心臟收縮時,動脈把全身血液輸送到毛細(xì)血管。靜脈輸送血液返回心臟。Alsofunctioningincirculationisthelymphaticsystem.Someofthefluidthatsurroundscellsdoesnotreenterthebloodvesselsdirectly.Thisfluid,called
15、lymph,returnstotheheartbywayofanothersystemofchannels-thelymphvessels.Lymphnodesalongthesevesselsfilterthefluidbeforeitreenterstheblood.Thespleenisalargelymphaticorganthatfilterstheblood.淋巴系統(tǒng)也是循環(huán)系統(tǒng)的一個組成部分。一些細(xì)胞周圍的體液不是直接回流入血管通道,這種體液叫淋巴液,它是流經(jīng)另一個管道系統(tǒng)淋巴管而回流人心臟。沿淋巴管的淋巴結(jié)將淋巴液過濾,過濾后再回流人血液。脾是一個過濾血液的大淋巴器官。Ther
16、espiratorysystemtakesinoxygenfromtheairandexpelscarbondioxideandwatervapor.Airentersthenoseandmouthandtravelsthroughthelarynx,andtrachea.Thetracheadividestoentereachofthetwolungsandthendividesmorethan20timestoformaverylargenumberofsmallairspaces.Oxygenfromtheairentersthebloodthroughcapillariesinthew
17、allsoftheseairspaces,andthebloodreleasecarbondioxideintotheairspacestobeexhaled.呼吸系統(tǒng)從空氣中攝取氧氣,并將二氧化碳、水蒸氣排出體外??諝饨?jīng)鼻腔、口腔人喉管、氣管。氣管分成左右支氣管,各連結(jié)左右肺,左右支氣管再分枝20多次,在終端形成大量微小的肺泡。從空氣攝取的氧氣流經(jīng)這些肺泡壁內(nèi)的毛細(xì)血管流入血液,血液再經(jīng)肺泡把釋放出的二氧化碳排出體外。Thedigestivesystemconsistsofatubeextendingfromthemouthtotheanus.Init,foodandfluidsaretak
18、enin,movedthroughthebody,andbrokendownintosmallmoleculesthatareabsorbedintothecirculatorysystem.Thisbreakdown,knownasdigestion,isbothamechanicalandachemicalprocess.消化系統(tǒng)是一個從口腔直到肛門的管道。食物和液體在消化道里被吸收,在腸道里移動時,被分解成小分子物質(zhì)后再進(jìn)入循環(huán)系統(tǒng)。這種分解,即消化,是一個機(jī)械過程,也是一個化學(xué)過程。Foodentersthroughthemouth,wherechewingandsalivastart
19、tobreakitupandmakeiteasiertoswallow.Next,thefoodtravelsdownthroughtheesophagustothestomach.Contractionsofthestomachsmuscularwallcontinuetobreakdownthefoodmechanically,andchemicaldigestioncontinueswhenacidandenzymesaresecretedintothestomachcavity.食物進(jìn)入口腔里,咀嚼和唾液開始將食物粉碎,使之便于吞咽。接著,食物經(jīng)食管人胃。胃肌壁的收縮繼續(xù)機(jī)械化地分解食
20、物,而當(dāng)酸和酶分泌入胃腔時,化學(xué)性消化開始。Theliquifiedfoodgraduallypassesintothesmallintestine.Inthefirstpartofthesmallintestine,calledtheduodenum,enzymesfromthepancreasareadded.Theseenzymescompletethechemicalbreakdownofthefood.Thedigestionoffatisaidedbybile,whichismadeintheliverandstoredinthegallbladder.Thesmallintest
21、ineofanadultisabout21feet(6.4meters)long.Mostofitslengthisdevotedtoabsorbingthenutrientsreleasedduringthesedigestiveactivities.液體化食物逐漸進(jìn)入小腸。小腸的起始部分叫十二指腸,胰腺分泌的酶輔助食物消化。這些酶完成食物的化學(xué)分解。肝臟分泌的膽汁貯存在膽囊內(nèi),膽汁有助于脂肪消化。一個成年人的小腸有21英尺(6.4米)長。小腸的大部分腸段用來吸收消化過程中釋放的營養(yǎng)物質(zhì)。Theliquidremainderofthefoodentersthelargeintestine,o
22、rcolon,whichisabout12feet(3.7meters)long.Itismorethantwiceaswideasthesmallintestine.Inthelargeintestinemostofthefluidisabsorbed,andtherelativelydryresiduesareexpelled.液狀的剩余食物進(jìn)入大腸,或結(jié)腸,它大約有12英尺(3.7米)長。大腸是小腸的兩倍多寬。大部分液體在大腸內(nèi)被吸收,相對干化的殘余物被排出體外。Theurinarysystemmaintainsnormallevelsofwaterandofcertainsmallmo
23、leculessuchassodiumandpotassiuminthebody.Itdoesthisbypassingbloodthroughthekidneys,twoefficientfilteringorgansthatgetridofanyexcessofvariousmoleculesandconservethosemoleculesthatareinshortsupply.泌尿系統(tǒng)維持水分及體內(nèi)某些小分子物質(zhì),如鈉、鉀的正常水平。身體是通過讓腎過濾血液來做到這一點(diǎn)的。腎是兩個有效的過濾器官,它濾出各種多余的小分子物質(zhì),保留那些供應(yīng)不足的小分子物質(zhì)。Thefluidthatleav
24、esthekidneys,knownasurine,travelsthroughatubecalledtheuretertothebladder.Thebladderholdstheurineuntilitisvoidedfromthebodythroughanothertube,theurethra.從腎流出的液體,即尿,通過輸尿管人膀胱。膀胱起貯存尿液的作用,直到尿經(jīng)膀胱另一端的管道排出。Theendocrinesystem.Thetwosystemsthatcontrolbodyactivitiesaretheendocrinesystemandthenervoussystem.Thef
25、ormerexertsitscontrolbymeansofchemicalmessengerscalledhormones.Hormonesareproducedbyavarietyofendocrineglands,whichreleasethehormonesdirectlyintothebloodstream.內(nèi)分泌系統(tǒng)。內(nèi)分泌和神經(jīng)是調(diào)控機(jī)體活動的兩個系統(tǒng),前者依靠其化學(xué)信使激素發(fā)揮作用。激素是由各種內(nèi)分泌腺體制造,并直接被釋放入血流Amajorglandisthepituitary,whichislocatedunderthebraininthemiddleofthehead.It
26、producesatleasteighthormones,whichaffectgrowth,kidneyfunction,anddevelopmentofthesexorgans.Becausesomeofthepituitaryshormonesstimulateotherglandstoproducetheirownhormones,thepituitarycalledthemastergland.腦垂體是一個主要腺體,它位于頭中部腦下方。它至少分泌八種激素,這些激素對人體生長、肝功能及性器官發(fā)育有影響。因?yàn)槟X垂體分泌的一些激素促進(jìn)其他腺體分泌激素,所以腦垂體是主要腺體。Anotherg
27、land,thethyroid,islocatedbetweenthecollarbones.Itshormonecontrolstherateofthebodysmetabolism.Thesexorgans(ovariesendtestes)makethesexcellsandalsomakehormonesthatcontrolcertaincharacteristicsofmalesandfemales.Locatedontopofeachkidneyistheadrenalgland,whichproducescortisoneandadrenaline.Thepancreaspro
28、ducesnotonlydigestiveenzymesbutalso3insulinandglucagon,whichcontrolthebodysuseofsugarandstarches.另一個腺體,甲狀腺,位于鎖骨之間。甲狀腺激素調(diào)控著機(jī)體新陳代謝的速度。性器官(卵巢、睪丸)分泌性細(xì)胞和性激素,這些激素控制著男性和女性的某些特征。每邊腎上方是腎上腺,它分泌可的松和腎上腺激素。胰腺不僅分泌消化酶,而且分泌胰島素和高血糖素,這兩種激素控制機(jī)體的糖分及淀粉的消耗。Thenervoussystem.Thebrain,thespinalcordandthenerve-alsocontrolsbo
29、dyactivities.Thelowerpartsofthebraincontrolbasicfunctionssuchasbreathingandheartrateaswellasbodytemperature,hunger,andthirst.Abovetheseregionsarethecentersforsight,sound,touch,smell,andtaste,andtheregionsthatdirectvoluntarymuscularactivitiesofthearmsandlegs.Performedherearethehigherfunctionsofintegr
30、atingandprocessinginformation.神經(jīng)系統(tǒng)腦、脊髓及神經(jīng),也調(diào)控機(jī)體活動。腦的偏下部位控制著諸如呼吸、心跳、體溫、饑渴的基本活動。而腦的偏上部位則是視覺、聽覺、觸覺、嗅覺及味覺中心,也是指揮臂、腿隨意肌肉運(yùn)動的區(qū)域。神經(jīng)系統(tǒng)更高級的功能是整合、處理信息。Thebrainreceivesandsendsinformationbymeansofnerves,manyofwhichliepartlyinthespinalcord.Thespinalcordisprotectedbythespinalcolumn.Nervesenterandleavethespinalcor
31、dateachlevelofthebody,travelingtoandfromthearms,legs,andtrunk.Thesenervesbringinformationfromthevarioussenseorgans.Theinformationisprocessedbythebrain,andthenmessagesarecarriedbacktomusclesandglandsthroughoutthebody.腦通過神經(jīng)收集并傳送信息,許多神經(jīng)部分地分布在脊髓里。脊髓由脊柱保護(hù)。在機(jī)體每一級,神經(jīng)傳人、傳出脊髓,往返于臂、腿、軀體。這些神經(jīng)輸送來自各種感覺器官的信息。信息經(jīng)腦
32、處理后輸送回全身及腺體Thereproductivesystemisconstructeddifferentlyformalesandfemales.Themalereproductivesystemisresponsibleforproducing,transportingandmaintainingviablesperm(themalesexcell).Italsoproducesthemalesexhormone,testosterone,whichregulatesthedevelopmentofabeard,pubichair,adeepvoiceandotherbodilychar
33、acteristicsoftheadultmale.男、女性的生殖系統(tǒng)不同。男性生殖系統(tǒng)產(chǎn)生、輸送、維持能存活的精子(男性性細(xì)胞)。它也分泌男性激素、睪酮,以此調(diào)節(jié)胡須、陰毛、深沉嗓音極其他成年男子身體發(fā)育的特征。Thefemaleproductivesystemisresponsibleforproducingandtransportingova(thefemalesexcells),eliminatingovafromthebodywhentheyarenotfertilizedbysperm,nourishingandprovidingaplaceforgrowthofanembryow
34、henanovumisfertilizedbysperm,andnourishinganewbornchild.Thefemalereproductivesystemalsoproducesthefemalesexhormones,estrogenandprogesterone,whichregulatethedevelopmentofbreastsandotherbodilycharacteristicsofthematurefemale.女性生殖系統(tǒng)產(chǎn)生、輸送卵子(女性性細(xì)胞),將未受精的卵子排出體外,而當(dāng)精、卵結(jié)合時,女性生殖系統(tǒng)培養(yǎng)、提供胚胎生長場所,并孕育新生兒。女性生殖系統(tǒng)也分泌女
35、性性激素雌激素和孕酮,以此調(diào)節(jié)乳房及其他成熟女性身體發(fā)育的特征。Theskinisacompletelayerthatprotectstheinnerstructuresofthebody,anditisthelargestofthebodysorgans.Itkeepsoutforeignsubstancesandpreventsexcessivewaterevaporation.Thenervesintheskinprovidetactileinformation.Theskinalsohelpskeepthebodystemperaturecloseto37C,heatisconserv
36、edbyreducingbloodflowthroughtheskinorisexpendedbyincreasingbloodflowandbyevaporationofsweatfromtheskin.Hairandnailsareaccessorystructuresoftheskin.皮膚是保護(hù)肌體內(nèi)層結(jié)構(gòu)的完整層,也是機(jī)體的最大器官皮膚防御外來侵襲,防止過多水分蒸發(fā)。皮膚上的神經(jīng)提供觸覺信息。皮膚也能將體溫維持到98.6華氏度(約37攝氏度)。通過皮膚的血流量降低時,熱量就被儲存起來,通過皮膚的血流增加及汗液蒸發(fā)時,熱量就散發(fā)。頭發(fā)及指甲是皮膚的附屬結(jié)構(gòu)。CellsandTissue
37、s細(xì)胞與組織Inthispassage,youwilllearn:1.Humanbodysystemsasawhole2.Vitalpartsofacell3.FourkindsoftissuesCellsareorganizedintotissues,andtissuesarearrangedintoorgans,whichinturnaregroupedintosystems.Eachbodysystemservesitsspecificfunctions.Bearinmindhoweverthatthebodyfunctionsasawholenosystemisindependento
38、ftheothers.Theyworktogethertomaintainthebodysstateofinternalstability,termedhomeostasis.Nowletsbeginourdiscussionwithcells,thesmallestunitoflivingmatterthatcanexistbyitself.細(xì)胞構(gòu)成組織,組織構(gòu)成器官,器官又進(jìn)一步構(gòu)成系統(tǒng)。人體的每個系統(tǒng)都有其特定功能,但是,請記住這里:機(jī)體是作為一個整體來發(fā)揮作用的,沒有哪個系統(tǒng)能夠獨(dú)立于其他系統(tǒng)而存在,是整體系統(tǒng)共同作用保持了機(jī)體內(nèi)部的穩(wěn)定狀態(tài),即體內(nèi)平衡。現(xiàn)在,我們先討論細(xì)胞-能獨(dú)立存
39、在活性物質(zhì)的最小單位。Thebodycanbestudiedfromitssimplesttoitsmostcomplexlevel,beginningwiththecell.Allbodyfunctionsresultfromtheactivitiesofbillionsofthespecializedcells.Someplantsandanimalsconsistofonlyasinglecell.Othersarecomposedofmanybillionsofcells.從細(xì)胞開始,我們能夠從最簡單到最復(fù)雜的水平來研究機(jī)體。所有人體的功能都來自億萬個特定細(xì)胞活動有的動植物僅由一個細(xì)胞
40、構(gòu)成,其他生物則由億萬個細(xì)胞構(gòu)成。Cellsexistinavarietyofshapesandsizes.Theymay,forexample,becube-shapedorflat.Scientistswhostudycellshavedeterminedthatasinglecellmaybeaslargeasatennisballorsosmallthatthousandswouldfitonthepointofaneedle.Theyolkofahenseggisactuallyaverylargecell.Bycontrast,bacteriaeachoneofwhichisati
41、nycellareamongthesmallestcells.Regardlessofitsshapeorsize,everycellcontainsthe“machinery”neededtomaintainlife.Whilenormallycellsfunctionwithgreatefficiency,theyaresubjecttovariousdisordersthatresultindisease.細(xì)胞有很多種形狀和大小。列如:有可能是立方形或扁形的??茖W(xué)家的研究發(fā)現(xiàn)單獨(dú)一個細(xì)胞可以像網(wǎng)球一樣的大小,或小到幾千個細(xì)胞才能剛好填滿針頭。雞的蛋黃是一個非常大的單細(xì)胞。微小的細(xì)菌呢算是
42、最小的細(xì)胞。不管是細(xì)胞大小或形狀的不同,每個細(xì)胞都有“需要存活的機(jī)制”。對每個正常的細(xì)胞,有效力的細(xì)胞,都存在許多問題,能導(dǎo)致疾病。Thesizeofcellsisusuallymeasuredinmicrons.Amicronisamillionthofameter,andabout25,000micronsequaloneinch.Thesmallestbacteriaareabout0.2micronindiameter.Theaveragecellinthehumanbodyabouttenmicronsindiameterisaspeckbarelyvisiblewithoutthe
43、aidofamicroscope.細(xì)胞的大小是以“微米”為長度單位。微米是一米的一百萬分之一,25000微米等于一寸。最小的細(xì)菌的圓徑是0.2微米。人體正常細(xì)胞的評價圓徑約十微米,肉眼勉強(qiáng)可以看得到,不需要顯微鏡的幫助。Thestudyofcellsisthebranchofbiologycalledcytology.Thesciencethatdealswithcellsonthesmalleststructuralandfunctionalleveliscalledmolecularbiology.研究關(guān)于“細(xì)胞”是生物學(xué)的分支,成為細(xì)胞生物學(xué)。在最微小水平針對細(xì)胞的結(jié)構(gòu)及功能的研究成為分
44、子生物學(xué)。Allcellsconsistofprotoplasm,the“l(fā)ivingjelly.”Theprotoplasmofatypicalcellformsthreevitalpartsthecellmembrane,thecytoplasm,andthenucleus.Themembraneenclosestheothercellstructures.Muchofthechemicalworkofthecellisdoneinthecytoplasm,whichsurroundsthenucleus.Thenucleus,enclosedbyitsownmembrane,isthec
45、ontrolcenterofthecell.(seeFigure1-1)所有細(xì)胞都是以“原生質(zhì)”,又稱“生命膠質(zhì)”所形成的。典型細(xì)胞的原生質(zhì)形成了三個重要部分,細(xì)胞膜、細(xì)胞質(zhì)和細(xì)胞核。細(xì)胞膜圈住了細(xì)胞的其他結(jié)構(gòu)。大部分的化學(xué)反應(yīng)都在細(xì)胞核周圍,細(xì)胞質(zhì)完成的。被自己的膜包圍的細(xì)胞核是細(xì)胞的控制中心。thecellsMembrane細(xì)胞膜Cellscansurviveonlyinaliquidmediumthatbringsinfoodandcarriesawaywaste.Forone-celledorganismsthisfluidisanexternalbodyofwatertheocean
46、,alake,orastream.Formany-celledplantsandanimals,however,themediumispartoftheorganisminplants,thesap;inanimals,theblood.細(xì)胞只能存活在能供養(yǎng)分又能排除廢物的液體媒介中。對單細(xì)胞生物來說,這就是細(xì)胞外的液體,如:大海里、河里或水流中。在對多細(xì)胞生物而言,這種媒介就是生物體的一部分,植物以汁液的形式,動物以血液的形式。Thecellmembraneissemipermeable,ordifferentiallypermeablesomesubstancescanpassthroug
47、hit,butotherscannot.Thischaracteristicenablesthecelltoadmitusefulsubstancesandtorejectharmfulsubstancesfromthesurroundingfluidaswellastoforceout,orexcretewasteproductsintothefluid.細(xì)胞膜是半滲透性或區(qū)別性的滲透膜,即某些物質(zhì)能滲透。這個特殊性使細(xì)胞對細(xì)胞周圍只吸收了需要的物質(zhì)而排除掉有害的物質(zhì),包括細(xì)胞內(nèi)的廢物通過液體化后排出細(xì)胞外。Thecellmembraneisanextremelythinbuttoughba
48、ndofproteinandphospholipidmolecules.Phospholipidsarechemicalssimilartostoredfat.Ontheevidenceofelectronmicrographs,biologistsbelievethatithasporesthroughwhichcertainsmallmoleculespassintact.Substancesprobablypassthroughthecellmembraneinseveralways.Smallchamberslocatedonthecellmembranecalledcaveolaeg
49、atherchemicalsignalsandchannelthemintothecells.Thecaveolaemayalsohelpdistributeproteinsthroughoutthecell.Largemoleculesenterthecellbyaprocesscalleddiffusion,inwhichtheymaybedissolvedbysubstancesinthecellmembrane.Theycanthenthroughthemembranewithoutdifficulty.Somecellstakeinlargemoleculesbymeansofpin
50、ocytosis.Inthisprocess,thecellmembraneformsapocketaroundlargemoleculesfloatingagainstit.Themolecule-and-fluid-filledpocketthenbreadsloosefromthemembranetobecomeabubblelikevacuole,andthevacuolethendriftsintothecytoplasm.Finally,thevacuolewallbreaksupandthemoleculesarereleasedintothecytoplasm.細(xì)胞膜是一種非常
51、薄但確以很堅(jiān)固的蛋白磷脂膜形成。磷脂是一種類似于儲存的脂肪的蛋白物質(zhì)。通過電子顯微鏡,生物學(xué)家相信細(xì)胞膜有孔,使較小的物質(zhì)完全可以通過。物質(zhì)可通過多種方式從細(xì)胞膜進(jìn)入。細(xì)胞膜上的結(jié)構(gòu),稱“小穴”,可接受化學(xué)信號再傳到細(xì)胞內(nèi)。小穴還可能參于細(xì)胞蛋白的分配。大的分子可通過“滲濾”進(jìn)入細(xì)胞內(nèi),或被細(xì)胞膜內(nèi)體分解,使它們更容易的進(jìn)入細(xì)胞內(nèi)。有些細(xì)胞可通過“胞飲”方式吞飲較大的物質(zhì)。這個過程是通過細(xì)胞膜在大物質(zhì)形成了“帶”把它包圍著,將它以泡狀的“液泡”引入細(xì)胞質(zhì),最后包圍解散,大物質(zhì)成功的進(jìn)入細(xì)胞內(nèi)。Thecytoplasm細(xì)胞質(zhì)Thecytoplasmismainlywater.Itswaterco
52、ntentvariesfromaminimumofabout65percenttoamaximumofabout95percent.Thesolidsinthecytoplasmincludegranularproteins,carbohydrates,dropletsoffat,andpigments.Thecytoplasmisthusacolloidalsystem.細(xì)胞質(zhì)主要成分是“水”。水分含量有百分之65到最高,百分之95。細(xì)胞質(zhì)內(nèi)的固體包括蛋白顆粒、碳水化合物、脂肪小滴和色素。細(xì)胞質(zhì)是膠體性的。Thecytoplasmmaybeeitherwateryorsyrupy,depen
53、dingontheconcentrationofsolidsdispersedinthefluid.Whenthesolidcontentdecreases,thesestructuresseemtovanish.Changesinconcentrationalsoproduceanapparentsteamingofthecytoplasmfromplacetoplacewithinthecell.Thisoccurs,forexample,whenfoodmoleculesenterthecell.細(xì)胞質(zhì)可能是水性或糖漿性,根據(jù)散在細(xì)胞質(zhì)內(nèi)固體的濃度。當(dāng)固體成分濃度增多,膜狀和纖維狀結(jié)果在
54、細(xì)胞質(zhì)顯出。而當(dāng)固體成分濃度減少時,膜狀和纖維狀的結(jié)構(gòu)就消失。濃度的變化還可以表現(xiàn)細(xì)胞質(zhì)內(nèi)局部流動的變化。食物分子進(jìn)入時可見。Mostofthecellsconstantworkofkeepingaliveisperformedinthecytoplasm.Herefoodmoleculesarechangedintothematerialneededforenergyandmaterialsforgrowthorstiffeningofthecellmembranearemanufacturesneededbytherestoftheorganism.Forexample,plantcell
55、scontainingchlorophyllmanufactureglucoseaplantfoodfromcarbondioxideandwater.大部分為了細(xì)胞存活的工作都在此處(細(xì)胞質(zhì))完成的。在此處,食物物質(zhì)將被轉(zhuǎn)化成提供細(xì)胞能量的材料和提供細(xì)胞生長的材料,或細(xì)胞膜的增強(qiáng)劑所生產(chǎn)。多細(xì)胞植物的細(xì)胞質(zhì)還是生產(chǎn)了對其他生物提供食物的物質(zhì)。列如:植物葉綠素所產(chǎn)生的葡萄糖,植物的食物,從水和二氧化碳合成。Oneofthecytoplasmskeyenergytransactionsoccursinthesausage-shapedmitochondria.Eachmitochondrionh
56、asaninnerandanoutermembrane.Likethecellmembrane,themembranesofthemitochondrionaresemipermeable.Foodmoleculesthatpassintothecytoplasmaretakenintothemitochondriaandoxidized,orburned,forenergy.細(xì)胞質(zhì)能量轉(zhuǎn)換的要點(diǎn)發(fā)生在錐狀形的線粒體。每個線粒體都有內(nèi)膜和外膜。像細(xì)胞膜一樣,線粒體的膜也是半滲透性。進(jìn)入線粒體的食物分子被氧化提供出能量。Theendoplasmicreticulum,anetworkofmemb
57、ranoustubes,runsthroughthecytoplasm.Intheopinionofsomebiologists,thisnetworkisacontinuousstructurethatbeginsatthecellmembrane,twiststhroughthecytoplasm,andendsatthemembranesurroundingthenucleus.Locatedalongtheendoplasmicreticulumaswellaselsewhereinthecytoplasmarenumerousribosomes.Thesetinygranulesco
58、nsistinpartofribonucleicacid(RNA).Proteinsaremanufacturedattheribosomes.TheGolgicomplex,orGolgiapparatus,isamembranousstructurecomposedofstacksofthinsacs.NewlymadeproteinsmovefromtheendoplasmicreticulumtotheGolgicomplex,wheretheyarestoredforlatersecretion.胞漿網(wǎng),在細(xì)胞質(zhì)中由膜所形成管狀的網(wǎng)。有些生物學(xué)家認(rèn)為這是一種連續(xù)性結(jié)構(gòu),始點(diǎn)為細(xì)胞膜,
59、在細(xì)胞膜內(nèi)扭絞,終點(diǎn)為細(xì)胞核的膜。分布于胞漿網(wǎng)包括細(xì)胞質(zhì)中某處的是核糖體。這些小顆粒是核糖核酸所形成的。蛋白質(zhì)是在核糖體合成的。高爾基體是膜狀囊包堆成層形成的。新合成的蛋白從胞漿網(wǎng)移動到高爾基體儲存,為了后來分泌所用。Vacuolesdriftthroughthecytoplasmandusuallycarryfoodmoleculesinsolution.Lysosomesarestructuressomewhatsimilarinappearancetovacuolesbutdenser.Theyappeartohaveadigestivefunction.Eachlysosomeisfi
60、lledwithdigestiveenzymesandencasedinamembrane.Lysosomesarebelievedtobreakdownfoodsubstancesbroughtintothecellbypinocytosis.IthasbeensuggestedthattheGolgicomplexplaysapartintheformationoflysosomes.在細(xì)胞質(zhì)內(nèi)游動的液泡一般是帶著食物分子的溶液。溶酶體是一種有點(diǎn)像液泡的結(jié)構(gòu),但較稠。它是用來消化的。每個溶酶體的膜內(nèi)都是添裝著消化酶。我們認(rèn)為它是消化通過“胞飲”方式的大物質(zhì)。也認(rèn)為高爾基體對溶酶體的形成有關(guān)
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